David Cox stated the game is a reboot of the franchise and is not a part of the original series canon. The setting of Lords of Shadow is during "the end of days" in the year of 1047. The Earth's alliance with the Heavens has been threatened by a malevolent force known as the Lords of Shadow. A dark spell has stopped the souls of the deceased from leaving, while evil creatures inhabit the dying land and attack living people.
The main character, Gabriel Belmont (voiced by Robert Carlyle), is a member of the Brotherhood of Light, an elite group of holy knights who protect and defend innocent people against the supernatural creatures. Gabriel's wife Marie (voiced by Natascha McElhone) was brutally murdered by one of them, and her soul cannot leave as it is trapped in limbo. Because she is now neither alive nor dead, she realizes what is at stake and guides Gabriel to his destiny to save the world as he investigates the dark spell. He travels the destroyed land, meeting other characters, such as the oldest living member of his order, Zobek (voiced by actor Sir Patrick Stewart, who additionally narrates the game). Two masks referred to as the God and Devil Masks lie at the centre of the plot, with the God Mask having powers to resurrect the dead. Gabriel intends to defeat the three factions of the Lords of Shadow in order to obtain the pieces of the God Mask and bring back his deceased wife.
Gabriel is sent by the Brotherhood of Light to the Lake of Oblivion, where his deceased wife, Marie, tells him that Spirits who founded the Brotherhood said that the Lords of Shadow's power will save the world. A man from the Brotherhood called Zobek introduces himself to Gabriel, who states that a prophecy has been kept a secret by a select few, which tells of a pure-hearted warrior who will claim the Lords of Shadow's power to overcome evil. He says that they must enter the lands of the Dark Lords in order to unite the Heavens with the world again, and that with this Gabriel can bring Marie back from the dead. Gabriel begins his journey at the Land of Lycans, where he encounters the Dark Lord, Cornell (Richard Ridings), who tells Gabriel that the three founding members of the Order fought the spawns of Satan in God's favour, and in their commitment to the crusade they found three places where God's power was intense to transform into the Spirits with power only second to God's. After they ascended to the Heavens they left behind their dark sides, the Lords of Shadow, which included Cornell. After defeating Cornell, Gabriel receives a piece of the God Mask before travelling to the Land of the Vampires.
Gabriel enters the castle of the Vampire Dark Lord, Carmilla (Sally Knyvette), where meets her daughter, Laura (Grace Vance), who decides to spare Gabriel after witnessing the love between him and Marie's spirit. A battle ensues between Gabriel and Carmilla, where he kills her for the second piece of the God Mask. Gabriel then departs for the Land of the Necromancers for the last part of the mask. Zobek then appears before Gabriel with the Devil Mask over his face, and divulges that he is the Lord of the Necromancers and that he grew tired of the Lords of Shadow diverting the power amongst the three of them instead of combining it. Orchestrating the events of the story, he searched Hell for this power until an evil force entered him and expanded his knowledge of the dark arts, which allowed him to cast the spell that separated the Earth from the Heavens so that the Spirits would contact the Brotherhood. Zobek discloses he used the Devil Mask on Gabriel to kill Marie and that all he needed was for Gabriel to restore the power of the Spirits to avoid suspicion from them.Satan (Jason Isaacs) emerges and takes the God Mask from Zobek, revealing himself as the mastermind who gave Zobek his powers so he could have revenge on God and return to the Heavens.Gabriel confronts Satan and defeats him, releasing souls of the deceased from limbo. Gabriel discovers the God Mask cannot bring Marie back and that it only allows him to see through God's eyes. Marie tells him he has been given a new life to redeem himself before she departs with the God Mask.
The story is expanded in two DLC packs titled Reverie and Resurrection and both episodes take place in between the ending and the epilogue. Reverie has Gabriel returning to Carmilla's castle to contain an ancient evil, the Forgotten One, with the help of Laura who is also a playable character that assists Gabriel in some sequences of the DLC. Before entering a portal, Laura tells Gabriel he cannot enter it in his mortal form and dies after she has Gabriel drink her blood, which turns him into a vampire. During Resurrection, the Forgotten One aspires to destroy the humans' world, but he is defeated by Gabriel who claims his power for himself. Corrupted by the Forgotten One's power, Gabriel destroys his Combat Cross and leaves through the portal. In a post credits scene after the main game's ending, Zobek is seen alive during modern times, and has uncovered Gabriel living out an immortal life as a vampire who now calls himself Dracul. Zobek mentions the acolytes of Satan are preparing for his return and that they must stop him before he takes revenge on both of them. Before Gabriel disappears, Zobek tells him he will free him of his immortality if he helps him.
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